Cross-system problems routinely exceed the competence and jurisdiction of any single institution, yet coordination itself can become a new source of domination. This paper develops the concept of a polycentric constitutional ecology: a system of differentiated but connected institutions in which authority remains matched to function, evidence and claims can travel consequentially across boundaries, coordinating bodies remain contestable, and persistent failure can trigger institutional redesign. It integrates polycentric governance, functional legitimacy, boundary organizations, subsidiarity, metagovernance, experimentalist governance, democratic anchorage, and institutional reflexivity.