This paper investigates the reentrant alignment of neurobiosemiotic layers across the Kosmic Life-Function framework and reveals a deeper architecture of coherence-making in living systems. By analyzing the shifting roles of mitochondria, emotion, interoception, and cognition across ontologically irreducible life-functions, we expose life not as a machine of fixed parts but as a semiotic field of recursive constraint interpretation. Drawing on the insights of teleodynamics (Deacon), biosemiotics (Barbieri, Hoffmeyer), and evaluative emotion theory (Peil Kauffman), we show that function is not static but contextually reassigned in response to systemic coherence needs. Emotion emerges as a key integrative reweaver, modulating the alignment between energetic readiness, interpretive salience, and participatory meaning-making. The resulting teleodynamic gradient — from energetic substrate to participatory alignment — reveals life as a coherence-seeking intelligence, capable of reorganizing its own constraint syntax across scale. We conclude with implications for medicine, systems design, AI, and civic evolution, proposing a semiotic ontology of function and a new grammar of participatory becoming.
